Gramma's 138th birthday was celebrated at the San Diego Zoo in 2022. (San Diego Zoo/YouTube | Screenshot by NPR) No one knows exactly when Gramma the Galápagos tortoise was born on that volcanic chain ...

Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, died Thursday. She was about 141. The zoo’s care team “made the compassionate and exceptionally difficult ...

Gramma, an estimated 141-year-old Galápagos tortoise, died Thursday at the San Diego Zoo, ending a near century-long chapter of what the zoo said was quiet, yet constant ambassadorship to generations ...

She was believed to be around 141 years old when she was euthanized due to "ongoing bone conditions related to advanced age," the zoo said Known as "the Queen of the Zoo," Gramma was in the first ...

Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, has died on Nov. 20. She was (about) 141. The zoo's care team "made the compassionate and exceptionally ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the Oval Office and there... How a 'sweet and shy' tortoise outlived empires and ...